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This occurs because hermaphrodites can outweigh losses in the female function by gaining paternity.

In an extended model we ask whether this favors the evolution of more male harm in hermaphrodites.

However, Charnov, assuming Bateman's principle, predicted that sexual conflict stemming from a preference for the male role would be important in hermaphrodite mating systems, creating a paradox.

Many hermaphrodite mating systems are based on conditional reciprocity with a preferred sexual role indicating sexual conflict.

Hermaphroditic animals often exhibit mating mechanisms that seem more damaging than those in species with separate sexes.

The great pond snail () seems to use a seminal product to manipulate its partner and mates in the male role when enough seminal fluid is available in the prostate gland.

Receipt of semen not only initiates egg laying in virgin animals, but also feminizes the mating partner later in life.